Principal Investigator

Photo of the Laboratory of Neurobiology of Learning and Memory principal investigator, David M. Smith, PhD.

David M. Smith, PhD

David M. Smith is a Professor of Psychology at Cornell University. He received his PhD in Psychology from The University of Illinois in 2001 under Michael Gabriel, and conducted postdoctoral research at the University of Washington with Sheri Mizumori. He is a recipient of the Stephen H. Weiss Junior Fellowship Teaching Award and heads The Laboratory of Neurobiology of Learning and Memory.

CV  |  Faculty Page | Contact


Current Members

Postdoc Hamid Turker

Hamid Turker, PhD
Postdoctoral Associate

Hamid studies the cognitive, computational, and neurobiological basis of how attention, memory, and control systems acquire structured knowledge about a semi-structured world and how that knowledge, subsequently, guides the deployment of those same systems. This is a tricky process given that these systems are limited and fallible. To investigate how the mind nevertheless manages this to produce adaptive behavior, Hamid employs a range of tools: from fMRI, EEG, and pupillometry (in humans), to electrophysiology (in rodents), and computational modeling (in always-too-slow computers). He joined the Smith lab after receiving a B.Sc. and M.Sc. from Leiden University in the Netherlands and an M.A. and Ph.D. from Cornell University. In his spare time, he enjoys playing guitar. CV

Photo of current PhD student Julia (Dahae) Jun.

Julia (Dahae) Jun
Graduate Student

Julia is interested in the role of the prefrontal cortex in memory retrieval. She received her B.S. from Stony Brook University, where she worked with Dr. Ryan Parsons investigating neurobiological mechanisms of learning and memory using Pavlovian fear conditioning. Julia is currently investigating the differential role of the subregions of the medial prefrontal cortex in retrieving target memories and connections with other regions such as the anterior olfactory nucleus. She is using a variety of techniques, including chemogenetic and optogenetic manipulations.

Graduate student Emily Li (Shiping)

Emily Li (Shiping)
Graduate Student

Emily received her B.S. from the University of Illinois at Urbana Champaign and her M.S. from Boston University, both majors in Psychology. She is currently interested in studying the functions of retrosplenial cortex.

Photo of current PhD student Dev Laxman Subramanian.

Dev Laxman Subramanian
Graduate Student

Dev is studying the spatial information encoding properties in the hippocampus and the retrosplenial cortex. Specifically, he analyzes the electrophysiological data collected using micro-drive arrays from the rat brain during spatial navigation tasks in a couple of different environments (Open field box foraging & continuous T-maze spatial alternation). His analyses are focused on understanding the similarities and differences in the spatial encoding properties in the hippocampus and the retrosplenial cortex at both individual neuron and population activity levels. He hopes that understanding the various methods of information encoding properties that the brain uses will provide us with insights into the neural algorithms that can have potential uses in advancing Artificial intelligence. In his free time, Dev likes to hike, spend time in nature, watch sports, and plays cricket for Cornell. CV

Wendy Yang
Graduate Student

Wendy is exploring the role that the anterior thalamus plays in hippocampal representations. She received her BS in Psychology from University of Illinois at Urbana-Champaign, where she worked with Dr. Justin Rhodes on the exercise-brain interactions. She is interested in learning about how various neural substrates are involved in the learning and memory circuits.

Photo of current lab research technician Anthony Coffin-Schmitt.

Anthony Coffin-Schmitt
Research Technician

Anthony assists Dr. Smith and other lab members with experiments, building microdrives and other devices for neurophysiological recordings, and rodent care and training. He previously worked in a bioinformatics research lab at UMass Medical School. Outside the lab, he enjoys going on hikes, exploring around Ithaca, and spending time with family.

Undergraduate student researcher Seychelle Balog.

Seychelle Balog
Undergraduate Research Assistant

Seychelle is a senior majoring in Biology and Society and minoring in Psychology and East Asian Studies. She is interested in studying the retrosplenial cortex and its role in contextual memory. Outside of the lab, she is a captain of the varsity women’s tennis team and a member of Rho Psi Eta.

Undergraduate student researcher Sarah Caldwell

Sarah Caldwell
Undergraduate Research Assistant

Sarah is a Human Development major minoring in Inequalities Studies on the Health Equity track. She is interested in working with adolescents to explore how social environments calibrate different neural mechanisms and the implications this can have on later risks for various psychopathologies.

Photo of undergraduate student research Emily Cheng.

Emily Cheng
Undergraduate Research Assistant

Emily is a sophomore majoring in statistical science and minoring in psychology. She is currently studying the role of the anterior thalamus in context-dependent memory. Outside of the lab, she is a part of Cornell’s taekwondo team and Alpha Chi Omega.

Undergraduate student researcher Angela Choi.

Angela Choi
Undergraduate Research Assistant

Angela is a junior studying Biological Sciences with a concentration in Human Nutrition. She is interested in the role of the medial prefrontal cortex in memory retrieval. Outside of the lab, she enjoys playing video games and volunteering at hospice centers.

Photo of lab member Molly Flanagan.

Molly Flanagan
Undergraduate Research Assistant

Molly is a junior studying Biological Sciences with a concentration in Animal Physiology. She is interested in memory retrieval, decision making, and the neural mapping of the brain. Outside of the lab, she enjoys volunteering her time to work with both small animals and dairy cattle, and is an active member of several club sports teams on campus.

Photo of lab member Jeffrey Ho.

Jeffrey Ho
Undergraduate Research Assistant

Jeffrey is a sophomore studying human development with a minor in business. He is fascinated by the base neurobiological and neurochemical interactions that inform higher-level processes such as memory and thinking. He is specifically interested in the neuronal mechanisms by which memory is represented. Outside the lab, he is involved in Red Cross, iGEM and Cornell Healthcare Review.

Undergraduate student researcher Alice Jang.

Alice Jang
Undergraduate Research Assistant

Alice is a sophomore studying Biological Sciences with a concentration in Neurobiology and Behavior. Her research interests surround the neurophysiological basis of memory and is currently assisting graduate student Julia with her project investigating the role of the medial prefrontal cortex and its sub-regions in modulating memory retrieval. Outside the lab, Alice is a part of the Brain Exercise Initiative, Cornell Healthcare Review, Cornell Undergraduate Research Journal, and The Women’s Network and spends her free time watching movies and playing The New York Times games.

Undergraduate student researcher Claire Jiang.

Claire Jiang
Undergraduate Research Assistant

Claire is a junior majoring in Animal Science and minoring in psychology. She’s developed an interest in understanding the intricate higher-level processes of the brain involved with memory and learning as well as understanding the biological mechanisms of the brain. Outside the lab, she loves to read and is a part of GlobeMed, PATCH, and the Cornell Surgical Society.

Undergraduate researcher Timothy Johnson

Timothy Johnson
Undergraduate Research Assistant

Tim is a sophomore studying Cognitive Science. He is interested in the mechanisms by which memories are encoded and represented. Outside of the lab, he is involved in the Cornell Undergraduate Research Board, Cornell Healthcare Review, and Cornell Undergraduate Research Journal.

Undergraduate student researcher Irene Liang.

Irene Liang
Undergraduate Research Assistant

Irene is a rising junior double majoring in Cognitive Science and Psychology. She is deeply interested in pursuing interdisciplinary approaches to systems neuroscience, specifically in regard to understanding the neural circuitry of memory retrieval and learning. In her free time, Irene enjoys volunteering as a TED Translator and trying new things—whether that is recipes, water sports, or arts and crafts projects.

Undergraduate researcher Andrea Pulido

Andrea Pulido
Undergraduate Research Assistant

Andrea is a senior studying Biological Sciences with a concentration in Neurobiology and Behavior. She is interested in memory retrieval, as well as the neural basis of learning and memory. Outside of the lab, she likes spending time with friends and family, long distance running, and volunteering at local centers. 

Undergraduate student researcher Hannah Schneider.

Hannah Schneider
Undergraduate Research Assistant

Hannah is a junior majoring in biological sciences (neurobiology and behavior) and minoring in psychology. She is interested in the interconnectedness of brain regions that give rise to memory and learning, and the role of behavior in this process. Outside the lab, she is involved in Alzheimer’s Buddies, The Steminist Movement, and intramural soccer and Spikeball.

Photo of lab member Izzy Wu.

Izzy Wu
Undergraduate Research Assistant

Izzy is a junior majoring in Psychology. She is interested in the neural basis of psychological disorders (especially generalized anxiety and depression) and its relationship with memory impairment. Outside of research, Izzy is involved in Cornell American Red Cross as well as a member of the Chinese Student Association. 


Lab Alumni

Former Postdoctoral Researchers

Photo of current Postdoctoral Associate lab member Wen-Yi Wu.

Wen-Yi Wu, PhD
Postdoctoral Associate and Research Associate – 2016-2023

Wen-Yi is interested in the neural basis of prosocial behavior, oxytocin and vasopressin. She investigates the neural mechanism of empathy-like prosocial behaviors of the rats and is also interested in the neurophysiological substrates of social and spatial representation in the rodent brain. She went on to a Assistant Professor position in the School of Medicine at Fu Jen Catholic University in Taiwan. Contact CV.

Photo of David Bulkin, PhD.

David Bulkin, PhD
Postdoctoral Associate – 2010-2015

Dave Bulkin is interested in how populations of neurons work together to support rich and complex behavior and hippocampal context representations. He was awarded the NARSAD Young Investigator Postdoctoral Fellow and went on as a Research Associate in the Melissa R. Warden laboratory at Cornell University. For more information, see his CV.

Photo of Lindsey Vedder, PhD.

Lindsey Vedder, PhD
Postdoctoral Associate – 2012-2014

Lindsey is a neurophysiologist interested how the hippocampus may interact with other brain regions for normal learning and memory processes. In the Smith lab, Lindsey investigated the relationship between the retrosplenial cortex and hippocampus during goal directed learning and memory. She went on to Binghamton University as a Postdoc and then as a Research Assistant Professor.

Photo of Patrick Gill, PhD.

Patrick Gill, PhD
Postdoctoral Associate – 2011

Patrick is an electrophysiologist and theorist interested in the ways neurons wire together to perform behaviorally-relevant tasks. He is especially interested in the neurocircuitry involved in the acquisition of expertise. As a post-doc in our lab, Patrick worked on multiple simultaneous tetrode recordings from the hippocampus and retrosplenial cortex of rats learning an auditory cued task, and also published work on Hippocampal Episode Fields (PDF).

Former Graduate Students

Former MS student Norma Hernandez.

Norma Hernandez
MS Student – 2019

Norma researched the anterior olfactory nucleus and ventral hippocampus roles in retrieval of contextually cued memory. She went on to be the lab manager of the Burwell Laboratory of Memory and Attention at Brown University. Thesis: Memory Retrieval Mechanisms in Context-dependent and Categorization Memory Tasks.

Photo of former PhD Student Adam Miller.

Adam Miller, PhD
Ph.D. Student – 2017

Adam researched systems-level changes during the learning process, and in the contributions of long-term memory to new learning. He went on as a Postdoc in the Frankland Lab at the University of Toronto. Thesis: The Role of the Retrosplenial Cortex in Spatial Cognition.

Former PhD student Greg Peters.

Gregory Peters, PhD
PhD Student – 2016

Greg examined the role of the medial prefrontal cortex and posterior cingulate cortex in a task already known to involve the hippocampus. Thesis: Contributions of the Medial Prefrontal Cortex to Managing Memory and Resolving Mnemonic Interference.

Former MS student Rachel Swanson.

Rachel Swanson
MS Student – 2015

Rachel conducted research on the functional contributions made by the retrosplenial cortex and the oscillatory co-activity across time in the hippocampus, retrosplenial cortex, and anterior thalamus. She went on to a Neuroscience PhD program at New York University. Thesis: Temporal and Spatial Coordination of the Hippocampus, Retrosplenial Cortex, and Anterior Thalamus.

Photo of former PhD student Matt Law.

Matthew Law, PhD
PhD Student – 2014

Matt graduated with a PhD after completing work on the limbic learning and memory system with a focus on the anterior thalamic nuclei. He went on as a Postdoc at the Barrow Neurological Institute at Phoenix Children’s Hospital. Thesis: Contextual Learning In The Limbic Memory Circuit.

Photo of former Master's student Orriana Sill.

Orriana Sill
MS Student – 2013

Orriana graduated with a masters degree after conducting a series of experiments on the role of the hippocampus in episodic memory. She went on to work as a data analyst. Thesis: Neurogenesis Dependent Learning And Implications For Episodic Memory In Rodent Models.

Former Undergraduate Researcher Assistants

Undergraduate student Jiarui (Ray) Fang

Jiarui (Ray) Fang
Undergraduate Research Assistant – 2024

Worked with post-doc Wen-Yi focusing on the role of the Anterior Olfactory Nucleus in contextually-cued odor discrimination. Ray completed an honors thesis titled “An Investigation into Anterior Olfactory Nucleus Neuronal Activity During Learning of a Complex Context Dependent Odor Memory Task”. He will be starting a PhD program at Boston University in the Dr. Michael Hasselmo lab.

Aditya Vinodh
Undergraduate Research Assistant – 2024

Worked with graduate student Julia Jun on infralimbic cortex and memory tasks and optogenetic experiments. Aditya is currently pursuing a Masters in Information Science degree at Cornell.

Undergraduate student Lizbeth Genao

Lizbeth Genao
Undergraduate Research Assistant – 2023

Liz investigated the medial prefrontal role in memory retrieval using a variety of behavioral tasks. She is currently a Medical Assistant at Manhattan Restorative Health Clinics and plans to pursue a medical degree.

Undergraduate student researcher Lilyanna Gross.

Lilyanna Gross
Undergraduate Research Assistant – 2023

Lily investigated the role of the anterior thalamus in spatial learning tasks and how the disruption of anterior thalamus activity affects retrosplenial cortex firing. She is currently a NIH Post-baccalaureate IRTA Fellow with Dr. Patti Brennan in Advanced Visualization Branch.

Photo of undergraduate student Alex Hodder.

Alexandra Hodder
Undergraduate Research Assistant – 2023

Alex investigated the effect of chemogenetic inactivation of the anterior thalamus on spatial learning using behavioral tasks such as T-maze alternation and Morris water maze. She also assisted with the electrophysiology recording of the dorsal hippocampus when the anterior thalamus activity was disrupted. Currently plans on pursuing a medical degree.

Photo of undergraduate student Ben Lederman.

Benjamin Lederman
Undergraduate Research Assistant – 2023

Ben worked with post-doc Wen-Yi on odor interference tasks, optogenetic experiments, and coding behavioral data. He went on as a MD student at the Keck School of Medicine of USC.

Photo of lab member Audrey Liu.

Audrey Liu
Undergraduate Research Assistant – 2023

Audrey worked with graduate student Julia Jun and ran odor discrimination task experiments.

Undergraduate student Aerin Mok

Aerin Mok
Undergraduate Research Assistant – 2023

Aerin worked with graduate student Julia Jun and ran odor discrimination tasks and DREADD experiments to activate or suppress neuronal firing in prelimbic and infralimbic cortexes.

Photo of undergraduate student Rebecca Shannon.

Rebecca Shannon
Undergraduate Research Assistant – 2023

Rebecca completed an honors thesis titled “Sex Differences in the Role of the Infralimbic Subregion of the Medial Prefrontal Cortex in Modulating Memory Retrieval” and plans on pursuing a medical degree.

Undergraduate student Sam Vucic

Sam Vucic
Undergraduate Research Assistant – 2023

Sam implementing animal models determine relationships between brain regions, brain functions, and behavior. His is currently a NIMH Post-baccalaureate IRTA Fellow with Dr. Tonya White in Social and Cognitive Developmental Neuroscience.

Photo of current undergraduate student Sabrina Giaimo.

Sabrina Giaimo
Undergraduate Research Assistant – 2022

Sabrina worked with graduate student Julia Jun, investigating the role of the prefrontal cortex. She plans on pursuing a PhD in clinical psychology.

Lab alumni Zichen (Cleo) He

Zichen (Cleo) He
Undergraduate Research Assistant – 2022

Zichen (Cleo) worked on the comparative roles of the retrosplenial cortex, the hippocampus, and the anterior thalamus in spatial learning and memorization. She went on to a Psychology & Neuroscience Ph.D. program at Duke University.

Megan Spurney
Undergraduate Research Assistant – 2021

Megan investigated the anterior nucleus of thalamus using chemogenetics. She went on to work at the NIH Intramural Research Program with a team studying fMRI and plans on applying to graduate programs.

Photo of current undergraduate student Taylor Vail.

Taylor Vail
Undergraduate Research Assistant – 2021

Taylor studied the anterior nucleus of thalamus with behavioral task experiments using optogenetics. She went on to work as a Psychiatric Research Assistant at Vanderbilt University Medical Center and plans on later attending medical school.

Photo of current undergraduate student Chiara Alvisi.

Chiara Alvisi
Undergraduate Research Assistant – 2020

Chiara investigated the ventral hippocampus and anterior olfactory nucleus in context and olfactory integration using optogenetics and chemogenetics. She went on to work as a healthcare consultant.

Photo of current undergraduate student Samantha Rabinovich.

Samantha Rabinovich
Undergraduate Research Assistant – 2020

Samantha assisted graduate students on their research. She plans on investigating Alzheimer’s disease before attending medical school.

Photo of current undergraduate student Kimaya Raje.

Kimaya Raje
Undergraduate Research Assistant – 2020

Kimaya completed her honors thesis titled, “The Role of the Ventral Hippocampus and the Anterior Olfactory Nucleus in Context and Olfactory Integration” and went on as a MD-PhD candidate at the University of Cincinnati.

Photo of current undergraduate student Eunice Yiu.

Eunice Yiu
Undergraduate Research Assistant – 2020

Eunice completed her honors thesis titled, “The Relationship between Spatial Occupancy Time and Firing Patterns of Hippocampal CA1 Neurons in Response to Changes in the Social Context”. She went on as a PhD student in Psychology at UC Berkeley.

Former undergraduate student researcher Pete Rigas.

Pete Rigas
Undergraduate Research Assistant – 2019

Pete worked on neurological data analysis, methods, and programming in the lab. He went on as a master’s student in Applied Statistics at Cornell University.

Former undergraduate student researcher Josh Dobbin.

Josh Dobbin
Undergraduate Research Assistant – 2017

Josh completed an honors thesis titled “The Role of the Ventral Hippocampus and Anterior Olfactory Nucleus in Contextually Cued Odor Discrimination” and was awarded the Magna Cum Laude level of honors. He went on to work as a research assistant at the Laboratory of Neuroendocrinology at The Rockefeller University.

Former undergraduate student researcher Betsy Hurtado.

Betsy Hurtado
Undergraduate Research Assistant – 2017

Betsy performed memory tasks with olfactory discrimination in various contexts to identify changes in neuronal response patterns and specifically associate them with learning impairments. She went on to work in clinical research at Memorial Sloan Kettering Cancer Cancer.

Former undergraduate student researacher Lindsay Rait.

Lindsay Rait
Undergraduate Research Assistant – 2017

Lindsay conducted experiments exploring the role of the hippocampus and olfactory system in the memory process with contextual odor discrimination tasks. She also mentored high school students interested in research. She went on to a Psychology PhD program at the University of Oregon.

Former undergraduate student researcher Sherry Shi.

Sherry Shi
Undergraduate Research Assistant – 2017

Sherry worked on a retrosplenial cortex tasks project with a graduate student.

Photo of former undergraduate student researcher Anna Serrichio.

Anna Serrichio
Undergraduate Research Assistant – 2016

Anna completed an honors thesis on the role of the retrosplenial cortex in spatial learning and navigation. She loves skiing and dancing. She may also love language as she went on to get a masters in Speech and Language Pathology from Columbia University.

Photo of former undergraduate student researcher Calvin Fang.

Calvin Fang
Undergraduate Research Assistant – 2015

Calvin completed an honors thesis on extracellular recordings in the prefrontal cortex of rats in relation to behavioral inhibition. He went on as a NIH Postbac. Intramural Research Training Award (IRTA) program and is now a MD-PhD candidate at Yale University focusing in Neuroscience.

Former undergraduate student researcher Jacob Frith

Jacob Frith
Undergraduate Research Assistant – 2015

Jacob was a Human Development major concentrating in behavioral neuroscience and went on to work in business development in Boston.

Former undergraduate student researcher Marc Harrison.

Marc Harrison
Undergraduate Research Assistant – 2015

Marc was listed as a co-author on a paper looking at retrosplenial cortical neuron encoding. He went on to work as a research assistant at the USC Laboratory of Neuroimaging and is now a Psychology PhD student at Stanford University.

Photo of former undergraduate student researcher Jason Li.

Jason Li
Undergraduate Research Assistant – 2015

Jason completed an honors thesis on the role of the RSC during a delay memory task. He is off doing less important things now like studying medicine at Georgetown University.

Former undergraduate student researcher Sophia Ramos.

Sophia Ramos
Undergraduate Research Assistant – 2015

Sophia was a Human Development major with a double concentration on human neuroscience and personality development. She went on to medical school at UCLA.

Former undergraduate student researcher David (Yeunhyung) Yu.

David (Keunhyung) Yu
Undergraduate Research Assistant – 2015

David co-authored three poster presentations while working with graduate student Adam Miller on retrosplenial cortical neural populations and spatial memory. He went on to a MPH at Columbia University.

Photo of former undergraduate student researcher Cara Bekowitz.

Cara Berkowitz
Undergraduate Research Assistant – 2014

Cara completed an honors thesis on the distinct roles of the infralimbic and prelimbic regions of the medial prefrontal cortex in support of memory retrieval. She went on medical school at Weill Cornell Medicine.

Former undergraduate student researcher Luke Grosvenor.

Luke Grosvenor
Undergraduate Research Assistant – 2014

Luke conducted an honors thesis on the role of the anterior thalamus in contextual fear learning. He went on as a NIH Intramural Research Training Award Fellow and is now in a mental health PhD program at John Hopkins University.

Photo of former undergraduate student researcher Madison Marcus.

Madison Marcus
Undergraduate Research Assistant – 2014

Madison completed an honors thesis investigating discrimination learning in the rat prefrontal cortex. An example of her published work can be downloaded here (PDF). She went on to law school at the University of Virginia.

Photo of former undergraduate student researcher William Mau.

William Mau
Undergraduate Research Assistant – 2014

William completed an honors thesis on the development of spatial and reward-related neural responses in the RSC during learning. He went on to pursue a PhD in Neuroscience at Boston University.

Former undergraduate student researcher Sarah Parauda.

Sarah Parauda
Undergraduate Research Assistant – 2014

Sarah investigating the role of the retrosplenial cortex input in support of hippocampal memory representations. She went on to medical school at the Weill Medical College of Cornell University and then a neurology residency.

Former undergraduate student researcher Athena Shea.

Athena Shea
Undergraduate Research Assistant – 2014

Athena conducted maze experiments to analyze roles of the hippocampus and retrosplenial cortex in spatial memory formation. She went on to work as an analyst and consultant in New York.

Photo of former undergraduate student researcher Carly Britton.

Carly Britton
Undergraduate Research Assistant – 2013

Carly completed an honors thesis and worked on a study investigating hippocampal population dynamics during an odor discrimination task. She went on to a masters in science writing at John Hopkins University.

Photo of former undergraduate student researcher Sam Levy.

Sam Levy
Undergraduate Research Assistant – 2013

Sam conducted early pilot experiments of a within-subject rodent model of the RIF phenomenon. He is currently in the Neuroscience PhD program at Boston University.

Photo of former undergraduate student researcher Leela Patel.

Leela Patel
Undergraduate Research Assistant – 2013

Leela investigated hippocampal population drift over time both within and between environmental contexts. She went on to volunteer with AmeriCorps and later attended Tufts Medical School.

Former undergraduate student researcher Jennifer Lieberman

Jennifer Lieberman
Undergraduate Research Assistant – 2012

Jennifer assisted on behavioral experiments studying contextual interference learning. She went on as Staff Research Associate at UC Davis and then as a lab manager at Columbia University.

Former undergraduate student researcher Jina Lim.

Jina Lim
Undergraduate Research Assistant – 2012

Jina researched the role of the anterior thalamus in a contextual-based odor list learning task with rats and examined the function of the anterior thalamus with temporary muscimol lesions. She went on to medical school at the University of Washington.

Former undergraduate student researcher Stephen McDowell.

Stephen McDowell
Undergraduate Research Assistant – 2012

Stephen worked with a graduate student study of cognitive deficits by examining brain lesions of rats to determine the functionality of the anterior thalamus in the greater memory circuit. He went on to pursue a MS in computer science at Cornell University.

Former undergraduate student researcher Maurice Petroccione.

Maurice Petroccione
Undergraduate Research Assistant – 2012

Maurice co-authored a paper titled “Hippocampal Context Process is Critical for Interference Free Recall of Odor Memories in Rats”. He went on to pursue a PhD in Biology at the Scimemi lab at SUNY Albany.

Photo of former undergraduate student researcher Jegath (Jiggy) Athilingam.

Jegath (Jiggy) Athilingam
Undergraduate Research Assistant – 2011

Jiggy completed an honors thesis on the role of the endocannabinoid system in learning-induced neuronal survival. She went on to a neuroscience PhD program at the University of California, San Francisco.

Photo of former undergraduate student researcher Rohini Bagrodia.

Rohini Bagrodia
Undergraduate Research Assistant – 2011

Rohini completed honors research examining immediate early gene expression in the anterior thalamus, retrosplenial cortex and the hippocampus during an olfactory learning task. Went on to assist with autism spectrum disorder research at UCLA and then a masters in clinical psychology at Columbia University.

Photo of former undergraduate student researcher Christopher David.

Christopher David
Undergraduate Research Assistant – 2011

Christopher conducted honors research on the role of the medial prefrontal cortex in olfactory discrimination learning. His thesis is titled “The Medial Prefrontal Cortex and Olfactory Learning”. Chris went on to the National Institute of Health in their Postbaccalaureate Intramural Research Training Award (IRTA) program and then medical school at University of Pittsburgh School of Medicine.

Photo of former undergraduate student researcher Grace Epstein.

Grace Epstein
Undergraduate Research Assistant – 2011

Grace completed an honors thesis investigating the role of the hippocampus in memory interference resolution and inhibition using a Retrieval-Induced Forgetting model. She went on to work as the Office Coordinator for Harlem Success Academy.

Photo of former undergraduate student Jade Wu.

Jade Wu
Undergraduate Research Assistant – 2011

Jade Wu received summa cum laude honors for her thesis on the role of the hippocampus in memory interference resolution and inhibition using a Retrieval-Induced Forgetting model and had a first author publication. Jade is now in a PhD program in clinical psychology at Boston University.

Former undergraduate student researcher Rachel Bavley.

Rachel Bavley
Undergraduate Student – 2010

Rachel conducted honors research on the contributions of hippocampal neurogenesis to cognitive functioning, culminating in a thesis titled “The Role of Hippocampal Neurogenesis in Context Discrimination”. She went on to the National Institute of Health in their Postbaccalaureate Intramural Research Training Award (IRTA) program and a MPH at Columbia University.

Former undergraduate student researcher Jess Masterton.

Jessica Masterton
Undergraduate Research Assistant – 2010

Jess co-authored a paper titled “Hippocampal Context Processing is Critical for Interference Free Recall of Odor Memories in Rats”. She went on to pursue a MFA in creative writing and a PhD in English and Comparative Literature at the University of Cincinnati.

Photo of former undergraduate student researcher Drew Melmed.

Drew Melmed
Undergraduate Research Assistant – 2010

Drew completed an honors thesis on the time-limited role of the retrosplenial cortex in discriminative learning. He went on to work as a research assistant at the Center for the Study of Traumatic Stress in Bethesda, MD.

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Yuliya Shteynberg
Undergraduate Research Assistant – 2010

Yuliya assisted on projects with training the rats and analyzing data. She went on as a MS student in Psychology at University of North Texas.

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Lena Germinario
Undergraduate Research Assistant – 2009

Lena analyzed temporal and spatial cues to episodic memory and assisted with research evaluating the role of the hippocampus in context-based odor discrimination tasks. She went on to law school at UC Hastings College of the Law.

Photo of former undergraduate student researcher Carolina Vega.

Carolina Vega
Undergraduate Research Assistant – 2009

Carolina worked on olfactory classification tasks. She went on to Tufts Medical School.

Photo of former undergraduate student researcher Ann Yang.

Ann Yang
Undergraduate Research Assistant – 2009

Ann acquired data on memory patterns affected by brain lesions and context interference cues in rats. She went on to a Masters of Science in Public Health (MSPH) degree in International Health, Health Systems at Johns Hopkins Bloomberg School of Public Health.

Photo of former undergraduate student researcher Dan Butterly.

Dan Butterly
Undergraduate Research Assistant – 2008

Dan helped develop our context based olfactory list learning task and conducted honors research on the role of the hippocampus in using contextual information to resolve interference. His thesis is titled “Hippocampal Lesions and the Importance of Context for Odor List Learning in Rats”. Dan is currently an analyst for an investment bank, probably making way more money than the rest of us.

Former Visiting Students

Marta Reales-Moreno
Visiting Graduate Student – 2023

Mara worked with graduate student Julia Jun on the role of the medial prefrontal cortex in cognitive processes like memory, including the use of optogenetic techniques. She received her Bachelor of Medicine and Surgery from the University of Girona and joined Smith’s lab as part of her doctoral program.

Former visiting student Daniela Betancurt.

Daniela Betancurt
Visiting Summer Research Assistant – 2018

Daniela recently finished a double major program in Microbiology and Biology with a minor in Psychobiology at the University of Los Andes in Bogota, Colombia.

Former visiting student Jenny Zaehringer.

Jenny Zähringer
Visiting Student – 2013

Jenny worked with graduate student Adam Miller investigating the role of the retrosplenial cortex input in support of hippocampal memory representations in rats. She went on to a PhD program at Central Institute of Mental Health in Mannheim, Germany.